Mike Conway will make his second IndyCar appearance of the year at this weekend's Detroit double-header after securing a deal to drive the second Dale Coyne Racing entry.
The Briton will be reunited with full-time Dale Coyne Racing driver Justin Wilson, who was his team-mate at Dreyer & Reinbold in 2010.
"I am very thankful for this opportunity with Dale and look forward to having a race weekend where I will have two chances to race with my fellow competitors," said Conway, who also raced for RLL at Long Beach.
"I miss being in the car full-time, and teaming with Justin again will be quite exciting for me."
Conway replaces Ana Beatriz, who was on a five-race deal. Team owner Dale Coyne paid tribute to the Brazilian while welcoming Conway into the team.
"First, I want to say what a great job Bia has done in the number 18 car," said Coyne.
"I think it was beneficial to run concurrent races with Bia where she could truly show her competitiveness as she did in Brazil and Indy.
"We are also excited to have a street course racer like Mike Conway join our efforts. Mike has shown over the years how competitive he can be on all types of circuits and we look forward to having our all-English team go at it at Belle Isle."
The team has not ruled out retaining Conway for future road and street events that do not clash with his WEC commitments, and also said that it is considering other drivers to fill the seat in the remaining races.
Candidates include Stefan Wilson, who tested with Coyne earlier in the year, and Oriol Servia, who is now available following Panther DRR's withdrawal from the championship after last weekend's Indy 500.