The new Rebellion Racing R-One World Endurance Championship LMP1 challenger ran for the first time at Paul Ricard on Wednesday.
The Toyota-engined coupe designed and built by ORECA was put through its paces by Swiss driver Mathias Beche in what was described as a "successful shakedown".
The roll-out follows a series of delays with the new car that mean Rebellion will rely on its old Lola-Toyota B12/60s for this weekend's WEC opener at Silverstone.
Rebellion team manager Bart Hayden said: "Today the Rebellion R-One has turned its first laps and we are absolutely delighted by that.
"Our mechanics have been in the ORECA factory for the past few weeks without a break, and so today's running of the car is a great reward for all the hard work."
It is planned that the R-One will make its debut at the second round of the WEC at Spa on May 3 after further testing at Ricard at the end of next week.
ORECA boss Hugues de Chaunac described the test of the car, his company's first completed ground-up LMP design, as a "proud moment".